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Evolution

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Evolution is a reoccurring mechanic in the Megami Tensei franchise, in which a demon transforms into another, typically more powerful demon.

Demon Evolutions

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Shin Megami Tensei III: Nocturne

Evolution Chart
Initial Level Evolved
Fairy Pixie 6 Fairy High Pixie
Night Lilim 12 Night Lilith
Beast Inugami 17 Avatar Makami
Snake Nozuchi 18 Dragon Xuanwu
Beast Nekomata 22 Holy Senri
Yoma Koppa Tengu 24 Yoma Karasu Tengu
Brute Momunofu 24 Vile Arahabaki
Yoma Dís 27 Wargod Valkyrie
Snake Naga 32 Snake Naga Raja
Snake Mizuchi 38 Dragon Qing Long
Yoma Ongkhot 42 Genma Hanuman
Fairy Setanta 48 Genma Cu Chulainn
Yoma Jinn 48 Yoma Efreet
Megami Sati 52 Lady Parvati
Yoma Pulukishi 53 Wargod Ganesha
Beast Suparna 58 Avian Garuda
Jirae Gogmagog 60 Entity Albion
Divine Throne 68 Seraph Uriel
Megami Scathach 69 Lady Skadi
Tyrant Abaddon 74 Tyrant Aciel
Tyrant Beelzebub (Human) 86 Tyrant Beelzebub (Fly)

Shin Megami Tensei IV

Evolution Chart
Initial Level Evolved
Holy Heqet 11 Lady Dzelarhons
Divine Angel 12 Divine Archangel
Jaki Gremlin 14 Wilder Raiju
Spirit Poltergeist 17 Spirit Quicksilver
Fairy Jack Frost 20 Jaki Wendigo
Yoma Koppa Tengu 22 Yoma Karasu Tengu
Beast Kabuso 23 Nymph Senri
Vermin Ubu 26 Jirae Tsuchigumo
Ghost Mou-Ryo 28 Spirit Inferno
Femme Shikome 31 Brute Yomotsu Ikusa
Night Incubus 37 Night Succubus
Undead Corpse 39 Fallen Nebiros
Fairy Kelpie 42 Yoma Valkyrie
Night Kaiwan 44 Fallen Decarabia
Jirae Kuebiko 49 Avatar Ukano Mitama
Drake Mushussu 51 Drake Tiamat
Avatar Chimera 52 Beast Orthrus
Holy Airavata 53 Jaki Girimekhala
Night Wild Hunt 58 Tyrant Abaddon
Vile Taotie 62 Wilder Taowu
Femme Dakini 69 Femme Kali

Shin Megami Tensei IV: Apocalypse

Evolution Chart
Initial Level Evolved
Divine Angel 16 Divine Archangel
Holy Heqet 17 Lady Dzelarhons
Yoma Koppa Tengu 20 Yoma Karasu Tengu
Spirit Poltergeist 23 Spirit Quicksilver
Fairy Jack Frost 26 Jaki Wendigo
Vermin Ubu 27 Jirae Tsuchigumo
Ghost Mou-Ryo 28 Spirit Inferno
Femme Shikome 34 Brute Yomotsu Ikusa
Fairy Kelpie 39 Yoma Valkyrie
Night Incubus 40 Night Succubus
Undead Corpse 42 Fallen Nebiros
Avatar Chimera 50 Beast Orthrus
Drake Mushussu 52 Drake Tiamat
Holy Airavata 57 Jaki Girimehkala
Night Wild Hunt 59 Tyrant Abaddon
Vile Taotie 62 Wilder Taowu
Femme Dakini 70 Femme Kali

Persona Evolutions

In the Persona games, the only Personas that are capable of Evolution are the ones used by party members. The protagonist's Personas do not actually evolve during the story of the games like the rest of the party members', but there are Personas that can be fused that are considered a Second Evolution.

Persona 3

Personas evolve on specific in-game dates. Party members do not have access to their full skill set before evolution, except for Koromaru and Shinjiro Aragaki, whose Personas never evolve, so they always have full access to their skill set.

Evolution Chart
Party Member Initial Persona Second Tier Persona Evolution Date
Yukari Takeba Io Isis November 5th
Junpei Iori Hermes Trismegistus November 22nd
Akihiko Sanada Polydeuces Caesar October 5th
Mitsuru Kirijo Penthesilea Artemisia November 18th
Fuuka Yamagishi Lucia Juno October 20th
Aigis Palladion Athena December 30th
Ken Amada Nemesis Kala-Nemi October 6th

Persona 3 Reload

Party members' skill sets are no longer restricted before they evolve as in prior releases of Persona 3. Instead, evolution grants them a second Theurgy. The exceptions are Koromaru, who starts with his second Theurgy, and Shinjiro Aragaki, who never evolves.

Persona 4

In Persona 4, evolution is referred to as transfiguration[1]. All Personas transfigure into their Second Tier Persona when their respective party member's Social Link has reached Rank 10.

Evolution Chart
Party Member Initial Persona Second Tier Persona
Yosuke Hanamura Jiraiya Susano-O
Chie Satonaka Tomoe Suzuka Gongen
Yukiko Amagi Konohana Sakuya Amaterasu
Kanji Tatsumi Take-Mikazuchi Rokuten-Maoh
Rise Kujikawa Himiko Kanzeon
Teddie Kintoki-Douji Kamui
Naoto Shirogane Sukuna-Hikona Yamato-Takeru

Persona 4 Golden

All Personas transfigure into their Second Tier Persona when their respective party member's Social Link has reached rank 10. Personas transfigure into their Third Tier Persona, officially referred to as an Ultimate Persona, once the protagonist does another Social Link event with each party member starting in January, only if the Second Tier Persona has been obtained.

Evolution Chart
Party Member Initial Persona Second Tier Persona Ultimate Persona
Yosuke Hanamura Jiraiya Susano-O Takehaya Susano-o
Chie Satonaka Tomoe Suzuka Gongen Haraedo-no-Okami
Yukiko Amagi Konohana Sakuya Amaterasu Sumeo-Okami
Kanji Tatsumi Take-Mikazuchi Rokuten-Maoh Takeji-Zaiten
Rise Kujikawa Himiko Kanzeon Kouzeon
Teddie Kintoki-Douji Kamui Kamui-Moshiri
Naoto Shirogane Sukuna-Hikona Yamato-Takeru Yamato-Sumeragi

Persona 5

All Personas evolve into their Second Tier Persona when their respective party member's Confidant has reached rank 10. Goro Akechi is the only party member whose Persona does not evolve.

Evolution Chart
Party Member Initial Persona Second Tier Persona
Morgana Zorro Mercurius
Ryuji Sakamoto Captain Kidd Seiten Taisei
Ann Takamaki Carmen Hecate
Yusuke Kitagawa Goemon Kamu Susano-o
Makoto Niijima Johanna Anat
Haru Okumura Milady Astarte
Futaba Sakura Necronomicon Prometheus

Persona 5 Royal

When viewing model sheets for the Personas in the Thieves Den, evolved Personas are referred to as "second evolution Personas" and "third evolution Personas"[2][3]. All Personas evolve into their second evolution when their respective party member's Confidant has reached rank 10. Personas evolve into their third evolution once the protagonist is approached by their respective party member in January, only if the second evolution has been obtained. The exception is Goro Akechi, who acquires his second evolution Persona automatically and can only acquire his third if the player has chosen the correct dialogue options while ranking up his Confidant and after securing the infiltration route to Shido's Palace.

Evolution Chart
Party Member Initial Persona Second Evolution Persona Third Evolution Persona
Morgana Zorro Mercurius Diego
Ryuji Sakamoto Captain Kidd Seiten Taisei William
Ann Takamaki Carmen Hecate Célestine
Yusuke Kitagawa Goemon Kamu Susano-o Gorokichi
Makoto Niijima Johanna Anat Agnes
Haru Okumura Milady Astarte Lucy
Futaba Sakura Necronomicon Prometheus Al Azif
Goro Akechi Robin Hood Loki Hereward
Sumire Yoshizawa Cendrillon Vanadis Ella
 
This section contains spoilers. (Skip Section)

Takuto Maruki's Persona, Azathoth, evolves into Adam Kadmon during his boss fight. Adam Kadmon is simply referred to as "Maruki's evolved Persona" in the Thieves Den Gallery.[4]

 
End of spoilers.

Persona 5 Strikers

The only character that undergoes evolution is Sophia, who transforms her Pithos into Pandora. Sophia receives an improved resistance, immediately learns Makougaon and gains access to her final skills, ala Persona 3.

Persona Q: Shadow of the Labyrinth

All Personas can evolve once both protagonists reach level 55, unlocking the "The Power of the Wild Card" request. After completing it, the protagonists' Personas will evolve immediately, while the other party members' will evolve upon reaching level 55.

Evolution grants each Persona their unique 4th skill. The exceptions are Zen and Rei, who are not Persona users, and Shinjiro Aragaki and Koromaru, whose Personas evolve only in terms of power, keeping their original forms.

Evolution Chart
Party Member Initial Persona Second Tier Persona Unique Skill
Makoto Yuki Orpheus Messiah Debilitate
Yukari Takeba Io Isis Tornado Boost
Junpei Iori Hermes Trismegistus Golden Gemini
Akihiko Sanada Polydeuces Caesar Conqueror Title
Mitsuru Kirijo Penthesilea Artemisia Punishment
Fuuka Yamagishi Lucia Juno Oracle
Aigis Palladion Athena Aegis Shield
Koromaru Cerberus Evolved Cerberus Deadly Vanguard
Ken Amada Nemesis Kala-Nemi Target Boost
Shinjiro Aragaki Castor Evolved Castor Whirlwind Swing
Yu Narukami Izanagi Izanagi-no-Okami Heat Riser
Yosuke Hanamura Jiraiya Susano-O Death Needle
Chie Satonaka Tomoe Suzuka Gongen Furious Fists
Yukiko Amagi Konohana Sakuya Amaterasu Inferno Boost
Kanji Tatsumi Take-Mikazuchi Rokuten-Maoh Uprising
Rise Kujikawa Himiko Kanzeon Into the Void
Teddie Kintoki-Douji Kamui Amrita
Naoto Shirogane Sukuna-Hikona Yamato-Takeru Powerhouse

Persona Q2: New Cinema Labyrinth

All Personas can evolve once the party obtains the 4 Sooty Films from requests 42 through 45 and defeat every Velvet Room attendant. After defeating Elizabeth, the protagonists' Personas will evolve immediately, while the other party members' will evolve upon reaching level 55, upon the next level up if they're already level 55, or immediately if they are level 99.

Evolution grants each Persona an improved version of their unique skill. The exceptions are Goro Akechi, Shinjiro Aragaki and Koromaru, whose Personas evolve only in terms of power, keeping their original forms, and the Navigators, whose evolutions instead reduce the cost of two of their skills by 1.

Evolution Chart
Party Member Initial Persona Second Tier Persona Unique Skill
Ren Amamiya Arsène Satanael Revolt Vanguard
Morgana Zorro Mercurius Trick Step
Ryuji Sakamoto Captain Kidd Seiten Taisei Revenge Style
Ann Takamaki Carmen Hecate Mistress Lash
Yusuke Kitagawa Goemon Kamu Susano-o Snack Expertise
Makoto Niijima Johanna Anat Unlimiter
Haru Okumura Milady Astarte Axe Bulwark
Futaba Sakura Necronomicon Prometheus -1 Cost: Zero Shift, Final Guard
Goro Akechi Robin Hood Evolved Robin Hood Detective's Ban
Yu Narukami Izanagi Izanagi-no-Okami Protecting Sword
Yosuke Hanamura Jiraiya Susano-O Kunai Dance
Chie Satonaka Tomoe Suzuka Gongen Counter Bash
Yukiko Amagi Konohana Sakuya Amaterasu Mirage Dance
Kanji Tatsumi Take-Mikazuchi Rokuten-Maoh Machismo Barrier
Rise Kujikawa Himiko Kanzeon -1 Cost: Encore, True Analysis
Teddie Kintoki-Douji Kamui Invincible Bod
Naoto Shirogane Sukuna-Hikona Yamato-Takeru Cost Reduction
Makoto Yuki Orpheus Orpheus Telos Judgment Card
Kotone Shiomi Orpheus Messiah Telepathic Bond
Yukari Takeba Io Isis Intercept Fire
Junpei Iori Hermes Trismegistus Called Shot
Akihiko Sanada Polydeuces Caesar Sonic Rush
Mitsuru Kirijo Penthesilea Artemisia Empress Pierce
Fuuka Yamagishi Lucia Juno -1 Cost: Healing Tide, Oracle
Aigis Palladion Athena Aegis Shielding
Koromaru Cerberus Evolved Cerberus Paradise Stroll
Ken Amada Nemesis Kala-Nemi Great Unity Link
Shinjiro Aragaki Castor Evolved Castor Effortless Style

Nomenclature

Names in Other Languages
Language Name Meaning
  Japanese 転生 Persona 4
Persona 4 Golden
Tensei		 Reincarnation

References

  1. ↑ "> Konohana Sakuya has transfigured into Amaterasu!" Persona 4 (Playstation 2). Developed by Atlus. Published by Atlus. English.
  2. ↑ "Model sheet of Sumire's second evolution Persona." Persona 5 Royal (Playstation 4). Developed by Atlus. Published by Atlus. English.
  3. ↑ "Model sheet of Sumire's third evolution Persona." Persona 5 Royal (Playstation 4). Developed by Atlus. Published by Atlus. English.
  4. ↑ "Model sheet of Maruki's evolved Persona." Persona 5 Royal (Playstation 4). Developed by Atlus. Published by Atlus. English.
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